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Position-effect variegation PEV when a gene normally in euchromatin is juxtaposed with heterochromatin by rearrangement or transposition. PEV is intensely studied in Escorts agency City of Milford using the white gene. Screens for dominant mutations that suppress or enhance white variegation have identified many conserved epigenetic factors, including the histone H3 lysine 9 methyltransferase SU VAR Genetic, molecular, and biochemical analysis of PEV in Drosophila has contributed many key findings concerning establishment and maintenance of heterochromatin with concomitant gene silencing. Genes that are abnormally juxtaposed with heterochromatin, either by rearrangement or transposition, show a variegating phenotype.
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